Family CALLIPALLENIDAE Hilton, 1942
Introduction
Monophyly of Callipallenidae was not recovered in morphological and total evidence analyses by Arango (2002) or Arango & Wheeler (2007).
The callipallenids are closely related to nymphonids, forming a strongly supported clade based on the presence of an oviger apophysis in males, and the development and type of larvae. However, the internal affinities of the Callipallenidae-Nymphonidae group remain to be tested, especially after inclusion of more callipallenid genera. Also, relationships of the Callipallenidae-Nymphonidae group with other pycnogonid lineages are not clear, as the support for the grouping with ammotheids is very low (Arango & Wheeler 2007).
Diagnosis
Chelifores present, scapes 1 or 2 segmented, chelae usually small, functional; palps either lacking, of 1-4 segments in males only, or only rudimentary knobs in both sexes; ovigers 6, 9 or 10-segmented, in both males and females, generally with compound spines. Legs long or short, with or without auxiliaries. Cement glands ventral.
General References
Arango C.P. & Wheeler W.C. 2007. Phylogeny of the sea spiders (Arthropoda; Pycnogonida) based on Direct Optimization of six loci and morphology. Cladistics 23: 255-293
Arango, C.P. 2002. Morphological phylogenetics of the sea spiders (Arthropoda: Pycnogonida). Organisms, Diversity and Evolution 2: 107-125
Hedgpeth, J.W. 1948. The Pycnogonida of the Western North Atlantic and the Caribbean. Proceedings of the United States National Museum 97(3216): 157-342, figs 4-53, charts 1-3
Hilton, W.A. 1942. Pantopoda (continued). II. Family Callipallenidae. Journal of Entomology and Zoology of Pomona College 34(2): 38-41
